Leaf senescence is determined by the maximum number of green leaves (GLFMX), the number of expanding and fully expanded leaves (EXPLF and TOTEXPLF, respectively). These variables are derived for each tiller cohort independently:

where LFMAX is a cultivar parameter expressing the maximum number of green leaves a healthy, adequately-watered plant will have after it is old enough to lose some leaves, SENESF is a species parameter representing the number of leaves per shoot senesced per 100 stress days, STDAYC is the estimated number of successive water stress days (see Abiotic stress section).

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